Forest Hill former News of the World editor Andy Coulson has denied ever being played a voicemail message left on James Bond actor Daniel Craig's phone by secret lover Sienna Miller.
The former Number 10 spin doctor refuted allegations from his one-time NotW colleague, journalist Dan Evans, that the editor was present when the reporter played the message about the Layer Cake co-stars.
And he denied ever being aware that convicted hacker Glenn Mulcaire - who had been employed by the now-defunct Sunday tabloid - was illegally listening in to voicemail messages.
He also hit out at the claim that he had "bullied" co-defendant and former colleague Clive Goodman.
The 46-year-old told the phone-hacking trial at the Old Bailey he was unaware that his own mobile was targeted by Mulcaire from 2006 until police informed him three years later.
Evans has already admitted conspiracy to hack phones at former employer the Sunday Mirror between February 2003 and January 2005, and the same offence at the News of the World between April 2004 and June 2010.
He has also pleaded guilty to conspiracy to commit misconduct in public office between January 2008 and June 2010, and perverting the course of justice by giving a false statement in High Court proceedings.
Evans, 38, has also claimed that Coulson knew about his activities and told him a voicemail he taped from Ms Miller to Mr Craig exposing their alleged affair was "brilliant".
Coulson, giving evidence for a fourth day, said Evans' account was wrong.
Defence counsel Timothy Langdale QC asked Coulson: "Dan Evans told the court that in September 2005 there was an occasion when he played to you a voicemail message left on Daniel Craig's telephone by Sienna Miller. Did any such incident take place?"
The defendant replied: "No, it did not."
Mr Langdale also asked if Coulson was aware of Mulcaire's hacking methods, to which the editor replied: "Absolutely not - no."
Coulson said there was no reason to believe hacking techniques were carried out because journalists had key contacts connected to celebrities, particularly in the case of the Miller-Craig affair.
Their identities were not disclosed in court, though it has already been reported that a relative of Ms Miller's then-boyfriend Jude Law had been supplying information about the British actor.
Coulson said the NotW's front page exclusive about Ms Miller and Mr Craig had come in the same week in 2005 when he had been busy overseeing a serialisation of British boxer Frank Bruno's biography.
He told the court: "It was a long, complicated book serialisation. I remember the publishers insisted on making changes quite late in the day, and I think the lawyers were involved as well.
"It took up a lot of my time."
Coulson said he believed information on the Miller-Craig affair may have come from Ms Miller's mother.
He added: "People in and around celebrities - their relatives, their agents, their PRs - will talk to newspapers."
He said he had asked staff working on the story not to refer to Mr Law as either a "sex addict" or a "love rat" in copy due to the paper's relationship with sources in his camp.
He said he also believed Mr Law was aware that some of those close to him were talking to the press.
Coulson said he could not remember having any specific dealings with Evans - who the tabloid recruited in January 2005 - during their time on the paper, but said they may have spoken.
He told the court: "I am sure we would have spoken to each other.
"If a new member of staff arrives I would normally make the point of saying hello and having a chat with them.
"I am sure that happened, but none (conversations) stick in my memory."
Coulson said his personal involvement as editor with stories and the reporters "varied".
He said: "There was no hard and fast set of rules in the way the story was researched and prepared."
Responding to suggestions earlier in the trial that he had bullied then-NotW royal editor Goodman by demoting him and reducing his responsibilities, Coulson said: "I'm not a bully.
"If I made a remark in conference or said something - it's a newspaper, that happens - I would apologise for it.
"I did nothing to undermine or attack Clive Goodman."
Coulson, of Wood Vale, is charged with conspiring to hack phones with Rebekah Brooks and former managing editor Stuart Kuttner, and conspiring with Goodman to commit misconduct in a public office. All seven defendants deny all charges against them.
Coulson told the court he "rubber-stamped" a payment from the NotW to a royal source for a story, despite being warned that being caught could end up with both parties in court.
The email exchange between Goodman and Coulson, in 2003, relates to payment for a "green book" of contacts for royal staff.
In the email, Goodman said he wanted to obtain the book for the "standard fee" of £1,000 from a royal policeman but the money should be paid in cash - something he needed Coulson to sign for.
The court heard Goodman said that getting caught making the payment meant the police officer "could end up on criminal charges, as could we".
Coulson, who said earlier that Goodman was prone to exaggeration and creating "unnecessary drama", said he signed off the payment.
He said: "I don't remember the email, but I have read them and thought a lot about them (in preparation for the trial).
"I didn't believe Clive was paying policemen, I still don't believe it.
"I think what I did was failed to address it properly with him. But I didn't (address it), I rubber-stamped it."
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